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- ItemOpen AccessSequence stratigraphic framework for top seal development: examples from the Skull Creek and Graneros shales, Denver basin(Colorado State University. Libraries, 1999) Edwards, Kimberly, K., author; Sutton, Sally J., advisor; Ethridge, Frank G., advisor; Almon, William R., committee memberIn general, the distal open marine shelf setting, typified by the Graneros Shale produces a rock with a greater and more uniform seal capacity relative to the rocks of a proximal open marine shelf setting, such as those of the Skull Creek Shale. A distal setting, which usually corresponds to the time of maximum transgression, may produce better seals because there is less coarse clastic sediment input, which allows slow deposition of clays from suspension to be the dominant depositional process. In this study, the higher capacity seal rocks occur in the upper parts of the TST, either within the condensed section or below it. The Skull Creek locations show seal occurrence to be stratigraphically higher on depositional topographic highs, and lower in areas that were topographically low at the time of deposition. Top seal capacity was quantified with mercury injection capillary pressure (MICP) analysis. Other physical characteristics of these marine shales were studied but only porosity, permeability, total clay, and hydrogen index consistently demonstrated a significant correlation with seal capacity in both units. Shales that are well laminated with a high percentage of total clay and/or total organic carbon with a type I-II (marine) kerogen may or may not qualify as the best seal. Top seal capacity may be more a function of rock fabric rather than mineralogy. For example, two samples may have exactly the same amount of quartz, as shown by XRD analysis, but thin section examination reveals that the majority of quartz in one sample is present as grains and in the other sample as cement. Samples with cement usually provide a better seal because they decrease the pore throat diameter, thus increasing the amount of hydrocarbons that can be trapped. Seal quality in both the Skull Creek and Graneros Shales is quite variable throughout each of the facies within the TST deposits.
- ItemOpen AccessInfluence of abiotic and biotic factors on the response of benthic macroinvertebrates to metals(Colorado State University. Libraries, 1995) Kiffney, Peter Michael, author; Clements, Will, advisor; Fausch, Kurt D., committee member; Kondratieff, Boris C., committee member; Chapman, Phillip, committee memberStream ecologists are well aware that chemical, biological, and physical characteristics of lotic systems vary spatially and temporally. With this in mind, I designed a series of experiments and field studies to examine the role of spatial variation in stream benthic macroinvertebrate communities in response to metals. Specifically, I tested the hypothesis that stream invertebrate communities from pristine streams of different size and altitude varied in their response to metals. To evaluate how metals affected biotic interactions, I manipulated invertebrate density, predation intensity, and metals in stream microcosms. Using stream invertebrate communities, I also designed an experiment and field survey to identify reliable bioindicators of metal contamination in western streams. Results from microcosm experiments and field studies showed that benthic invertebrate populations from high-altitude streams were more sensitive to the effects of metals than invertebrate populations from low-altitude streams. For example, Baetis sp. and Rhithrogena hageni from Little Beaver Creek (LBC), Colorado, (high-altitude stream) were significantly more sensitive to zinc than the same species from the South Fork of the Poudre River (SFP) (low-altitude stream) in stream microcosms. Results from field surveys showed that densities of most groups of aquatic insects (e.g., Ephemeroptera, Plecoptera, Trichoptera) were lower at high-altitude metal contaminated streams than those same groups at low-altitude streams. Other factors, such as variation in water temperature or nutrient concentrations between high- and low-altitude streams, could be lll responsible for these differences. However, because invertebrate responses were similar under controlled and field conditions, I hypothesize that differences in abundances between reference and contaminated locations was a result of metals. To determine if body size contributed to the variation in sensitivity of insects to metals, size measurements were made on species collected from LBC (high-altitude) and SFP (low-altitude). Measurements were also made on species from control and metal-treated stream microcosm. Most species were smaller at LBC (high-altitude stream) than the same species from SFP (low-altitude stream). For example, the mayfly Baetis tricaudatus and the caddisfly Arctopsyche grandis were significantly larger at SFP than LBC. In addition, insect body size was larger in metal-dosed microcosms than in controls. Brachycentrus sp., B. tricaudatus, R. hageni, Eohemerella infrequens, and P. badia were significantly larger in metal-treated microcosms than in controls. Logistic regression indicated survival in metal-dosed microcosms was less for small individuals than for larger individuals of the same species. These results suggest that some insect species from high-altitude streams were smaller than those from low-altitude streams, and that survival was greater for larger lifestages. Thus the variation in response of macroinvertebrates between different altitude streams observed in earlier studies may be due to differences in body size. The effects of low levels of metals (half the chronic levels of Cd, Cu, and Zn) on some species varied in relation to invertebrate density (low and high density) and invertebrate predation (no predators added and predators added). The abundance of Hydropsyche sp. was significantly lower in metal-dosed, high density treatments than in control, high density treatments. Moreover, the effects of an invertebrate predator (Hesperoperla pacifica) on Hydropsyche sp. was significantly greater in metal-dosed microcosms than in controls. These results suggest that metals interact with biotic factors to influence stream invertebrate community structure, and that effects occurred at metal concentrations lower than chronic criteria value. Toxicity experiments in stream microcosms showed that the abundance and species richness of aquatic insects were significantly reduced at 1x, 5x and 10x the United States Environmental Protection Agency chronic levels of cadmium, copper, and zinc (1x=1.1, 5.0, and 110 μg/L Cd, Cu, and Zn, respectively). Mayflies were the most sensitive group, as the abundance of Baetis sp. and Rithrogena hageni were significantly reduced in the Ix treatment. The response of Drunella grandis was size dependent, as small lifestages were significantly more sensitive than large lifestages. Stoneflies were also affected, but their response was more variable with abundances of some species (Pteronarcella badia) being reduced in the Ix treatment, whereas other species were unaffected (Sweltsa sp.). Heptageniid mayflies were consistently less abundant downstream of sources of metal contamination in the Arkansas and Eagle rivers, whereas the response of other measures were more variable. For instance, species richness and total density were greater at a metal-contaminated site on the Arkansas River compared with an upstream reference site. Therefore, results from this experiment and field survey suggest that changes in abundance of heptageniid mayflies may provide a reliable indicator of metal-contamination in western streams.
- ItemOpen Access5000 years at Crow's Roost in eastern Colorado(Colorado State University. Libraries, 1992) McDonald, Robert A., author; Jennings, Calvin H., advisor; Leyendecker, Liston, committee member; Zier, Christian, committee member; Theodoratus, Robert, committee memberMany areas on the high Plains of eastern Colorado have had little prehistoric research conducted on them. One of the areas where there is insufficient information about local prehistory is the area between the two major river drainages, the Arkansas and the Platte. Crow's Roost, a large sandstone cliff, located east of Colorado Springs, Colorado, is situated near this dividing line, known as the Palmer Divide. A series of prehistoric sites at Crow's Roost has been investigated over a period of five years by the Anthropology Department, University of Colorado at Colorado Springs. These investigations indicate that there has been prehistoric occupation at this locality dating back at least 5400 years. This study presents an interpretation of the culture history, and of recent paleoclimatic fluctuations for this locality.
- ItemOpen AccessThe bighorn sheep of Battlement Mesa - a low elevation population(Colorado State University. Libraries, 1991) Cunningham, Mary Louise, author; Bailey, James A., advisor; Alldredge, A. William, committee member; Lehner, Philip N., committee memberI studied bighorn sheep on Battlement Mesa, Colorado, from 12/88 to 7/90, to determine present and past herd status and distribution, so that 1) management options are clarified and 2) future management activities may be evaluated. I studied the herd by tracking 1 radio-collared ewe and obtaining visual observations and by systematically sampling study area units for sheep, sheep sign, potential competitors, potential predators and sheep carcasses. I recorded water sources that were encountered. I developed an historic perspective of the herd by searching agency files and local newspapers and interviewing local residents. The herd numbered up to 200-250 animals in the early 1900s and declined to an estimated 50 animals by 1970. Minimum herd sizes during 1989 and 1990 were 23 and 26, including lambs, respectively. Since 1961, the herd has abandoned approximately 56 km2 of historic range in the Mamm Peaks area. The herd decline corresponded with probable vegetation changes on Battlement Mesa, intensive livestock grazing through the 1950s, reports of poaching and an increasing elk herd. Sheep remained on the western portion of the range during winter and spring, 1989. Ewe/juvenile groups migrated to Anderson and Durant Gulches between 5/15-7/15/89 for lambing. Rams were more dispersed and in groups of 1-3, except during the rutting season. Ewe/juvenile group size ranged from 1-13. During dry months, (7/89, 8/89, 6/90) sheep concentrated in Anderson and Durant Gulches where a free-flowing spring and a seep were located. Bighorn sheep on Battlement Mesa appear limited by dense mountain shrub stands which separate all productive meadows from escape terrain and cover historic migration routes. A significantly greater use of the shale slope habitat contributed most to rejecting the null hypothesis that sheep use habitat types in proportion to their availabilities on Battlement Mesa. Sheep remained on shale slopes most (75% of all observations) of the time, using scattered grasses, forbs and shrubs for forage and seeps for water. Intensive and long term habitat management for bighorn sheep on Battlement Mesa is required. I suggest a 4-phase management program to improve existing range and later to reestablish and maintain historic migration corridors to productive historic summer range. Without management to improve existing conditions, this small, unique herd will remain static or decline.
- ItemOpen AccessFeed zone micromixing and its effect on continuous cultures of Saccharomyces cerevisiae(Colorado State University. Libraries, 1995) Mondani, Paul, author; Loftis, J. C., advisorInadequate mixing is known to be a common problem in the scale-up of bioprocesses, often leading to decreases in yield and productivity. To investigate the role of nutrient dispersion in continuous cultures, growth medium was fed into a laminar flow section of a loop that recirculates broth from a laboratory scale bioreactor. The intensity of micromixing at the feed site could be controlled by varying the axial distance a static mixer was placed upstream of the site. The intensity of the turbulent wake shed by the mixer was quantified by laser Doppler velocimetry and the Bourne dye reaction. By decreasing the size of the smallest turbulent eddy in the feed zone, less of the population is exposed to regions of either inadequate or excessive substrate concentrations. Yield vs. dilution rate curves were obtained through various mixing and feeding strategies. Reduced mixing was shown to delay the onset of the Crabtree effect and therefore improve the bioreactor's productivity.