Papilio. New Series
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Papilio (New Series), an entomology journal, is a scientific journal that covers the systematics and taxonomy and biology of butterflies, mostly from Colorado. There are about 700 species of butterflies in North America and about 270 in Colorado, New discoveries are made every year on the Colorado species. Systematics is the study of the kinds of butterflies that exist on our planet, and taxonomy involves the names of butterflies, including the description and naming of species new to science. Papilio (New Series) started in 1981 to name the butterflies in single publications. This digital collection includes the single issues as they are published.
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Browsing Papilio. New Series by Author "Spomer, Stephen M., author"
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Item Open Access Geographic variation and new taxa of western North American butterflies, especially from Colorado(Colorado State University. Libraries, 2008-12-03) Garhart, Matthew C., author; Stout, Todd, author; Kondla, Norbert G., author; Spomer, Stephen M., author; Fisher, Michael S., author; Scott, James A., author; Wright, David M., author; Marrone, Gary M., author; James A. Scott, publisherMichael Fisher is currently updating the 1957 book Colorado Butterflies, by F. Martin Brown, J. Donald Eff, and Bernard Rotger (Fisher 2005a, 2005b, 2006). This project has emphasized the necessity of naming certain butterflies in Colorado and vicinity that are distinctive, but currently have no name, as part of our goal of applying correct species/subspecies names to all Colorado butterflies. Eleven of those distinctive butterflies are named here, in the genera Anthocharis, Neominois, Asterocampa, Argynnis (Speyeria), Euphydryas, Lycaena, and Hesperia. New life histories are reported for species or subspecies of Neominois & Oeneis & Euphydryas & Lycaena that were recently described or recently elevated in status. Lycaena florus differs in hostplant, egg morphology, and somewhat in a seta on 1st-stage larvae. We also report the results of research elsewhere in North America that was needed to determine which of the current subspecies names should be applied to other butterflies in Colorado, in the genera Anthocharis, Neominois, Apodemia, Callophrys, Atlides, Euphilotes, PlebeJus, Polites, & Hylephila. This research has added additional species to the total of Colorado butterflies. Nomenclatural problems in Colorado Lycaena & Callophrys are settled with lectotypes and designations of type localities and two pending petitions to suppress toxotaxa. Difficulties with the ICZN Code in properly applying names to clines are explored, and new terminology is given to some necessary biological solutions.Item Open Access Speyeria hesperis and Speyeria atlantis are distinct species(Colorado State University. Libraries, 1998-02-20) Kondla, Norbert G., author; Scott, James A., author; Spomer, Stephen M., author; James A. Scott, publisherS. hesperis and S. atlantis are distinct species; past reports of "intergradation" between them actually represent polymorphism of silvering within S. hesperis. S. atlantis is always silvered, and has a darker (chocolate-brown to blackish-brown) unh disc; it occurs in eastern North America as S. a. atlantis, then ranges as S. a. hollandi over the boreal forest/aspen parklands from Man. To Alta., where it is widely sympatric with S. hesperis helena; S. a. hollandi also occurs in the Rocky Mts. in Alta.-B.C.-NE Wash.-N Ida., where it is sympatric with S. hesperis beani and S. hesperis brico (B.C., new subspecies). In the Black Hills, S. atlantis pahasapa (new subspecies) is sympatric with S. hesperis lurana. In S Wyo.-Colo.-N New Mex., S. atlantis sorocko (new subspecies) is sympatric with mostly-un silvered S. h. hesperis and mostly-silvered S. h. electa (=cornelia=nikias). S. hesperis has a redder unh disc, and ranges from Manitoba and the Black Hills westward to Alaska and the Pacific and south to New Mex.-Calif.; a majority (11 of 19) of its subspecies are usually-silvered, but only the subspecies in the extreme N and NE and S parts of its range are always silvered, and all subspecies across the middle of its range are predominantly unsilvered; silvered/unsilvered intergradation occurs within S. hesperis throughout the middle of the range, including the northern Sangre de Cristo Mts. of Colo. where unsilvered S. hesperis hesperis intergrades completely with silvered S. h. electa. Except in the extreme northern and southern ends of its range, older larvae of S. hesperis are blacker than larvae of S. atlantis.Item Open Access Taxonomic studies and new taxa of North American butterflies(Colorado State University. Libraries, 2006-04-28) Spomer, Stephen M., author; Guppy, Crispin S., author; Kohler, Steve, author; Kondla, Norbert G., author; Fisher, Michael S., author; Scott, James A., author; Schmidt, B. Chris, author; James A. Scott, publisherNew diversity is reported and discussed among North American butterflies. Several dozen new taxa are named. A new "sibling" species has been found to occur throughout the Rocky Mts., introducing a new butterfly species to most states in western U.S. and to southern Alberta and BC. Several taxa of Colias, Euphydryas, Lycaena, and Plebejus are raised to species status. Many name changes are made, and many taxa are switched between species to create several dozen new combinations. The relevance of species concepts to difficult groups of butterflies is explored.