C. P. Gillette Museum of Arthropod Diversity
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The C. P. Gillette Museum of Arthropod Diversity has its roots in the college museum established in the early 1890s, which was organized to support the teaching of zoology and entomology. Named for Clarence Preston Gillette (1859-1941), creator of the museum and the first professor of entomology at Colorado Agricultural College (now Colorado State University), the museum changed considerably over the 20th century. First housed in Old Main and curated by E. L. Burnett, the museum’s location changed several times before it was disbanded in 1947, at which point the insect collection became the property of the Department of Entomology. In the early 1990s, department staff named the collection the C.P. Gillette Museum of Arthropod Diversity. The museum currently holds approximately three million insect specimens and a noted reference library. The museum’s records contain two accession ledgers, the papers of butterfly collector Ray E. Stanford, newspapers, papers, and books of museum fellow Richard Holland.
Accession catalogs for insects and other arthropods dated 1890 to 1972 are found in the Gillette (C.P.) Museum of Arthropod Diversity archival collection. Many of the specimens were collected by C. P. Gillette, and entries typically include location, date, collector, specific habitats information, hosts, and associated other species.
These digital collections include publications, conference proceedings, and the newsletter Papilio. New Series.
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Item Open Access North American insect status review(Colorado State University. Libraries, 1975-1985) Perkins, Philip Don, compiler; Department of Entomology, Smithsonian Institution, authorItem Open Access New Papilionoidea and Hesperioidea from North America(Colorado State University. Libraries, 1981-11-25) Scott, James A., author; James A. Scott, publisherItem Open Access The life history and ecology of an Alpine relict, Boloria improba acrocnema (Lepidoptera: Nymphalidae), illustrating a new mathematical population census method(Colorado State University. Libraries, 1982-11-15) Scott, James A., author; James A. Scott, publisherThe egg, larva, and pupa of B. improba acrocnema are described and illustrated. The larval foodplant is Salix nivalis. There are five instars; overwintering occurs in the fourth instar, and perhaps also in the first instar. Adults fly slowly near the ground, and are very local. Males patrol to find females. Adults often visit flowers, and bask on dark soil by spreading the wings laterally. A binomial method is derived and used to determine daily population size. Daily population size may be as large as 655, and yearly population size may reach two thousand in a few hectares.Item Open Access The courtship of Phyciodes, and the relationship between Phyciodes tharos tharos and Phyciodes tharos morpheus (=pascoensis) in Colorado(Colorado State University. Libraries, 1986-03-05) Scott, James A., author; James A. Scott, publisherThe courtship of Phyciodes tharos/morpheus and P. campestris is described, and is very similar between species. Reared released female morpheus courted and mated readily with wild male tharos in Colorado. Together with the viability and fertility of the hybrids and the breakdown of the antennal and larval characters that distinguish them in W.Va.-Va. (where they behave as separate species), this evidence indicates that tharos/morpheus are not reproductively isolated in Colorado, where they seem to be a bit more than subspecies, but are not completely distinct species. Phyciodes have an interesting outcrossing mechanism involving timing of adult emergence, which ensures rapid spread of genes between parent forms after natural hybridization.Item Open Access Larval hostplant records for butterflies and skippers (mainly from western U. S.), with notes on their natural history(Colorado State University. Libraries, 1986-03-05) Scott, James A., author; James A. Scott, publisherLarval hostplants, based on observations of adult oviposition or larval feeding, are presented for butterflies (including skippers) from western United States (mostly Colorado), and many notes on egg placement, overwintering stage, behavior, and ecology of these species are given. A case of larval hostplant switching is reported in which Phyciodes picta (Nymphalidae) originally fed on Aster (Compositae) but now feeds on the recently introduced Convolvulus arvensis (Convolvulaceae), a completely unrelated weedy vine. Ovipositing females have distinctive slow hovering flights, but in Satyrinae these are not as distinct from normal flight. Females of Satyrinae, Speyeria, and Boloria oviposit rather haphazardly near the hostplants, and many Hesperiinae are somewhat haphazard about their choice of grasses/sedges or nearby plants for oviposition.Item Open Access Distribution of Caribbean butterflies(Colorado State University. Libraries, 1986-03-05) Scott, James A., author; James A. Scott, publisherItem Open Access Hostplant records for butterflies and skippers (mostly from Colorado) 1959-1992, with new life histories and notes on oviposition, immatures, and ecology(Colorado State University. Libraries, 1992-07-05) Scott, James A., author; James A. Scott, publisherHostplants of larvae, based on about 3090 records of observed ovipositions (a total of 1509) or discoveries of eggs, larvae, or pupae in nature, are presented for butterflies (including skippers), mostly from western United States, especially Colorado. The paper presents numerous new life histories, and many notes on egg placement, overwintering stage, behavior, and ecology. A new phenomenon of a butterfly egg mimicking a plant is reported, in which Nathalis iole eggs have changed color to match orange-yellow protruding foul-odor egg-shaped glands on Oyssodia papposa, presumably to benefit from lesser predation because predators think the eggs are inedible foul glands; this phenomenon is the complete opposite of the known cases of egg mimicry, in which plants produce structures mimicking butterfly eggs to deter oviposition (although possibly egg mimicry was the original origin of the precursor of the 0. oapposa glands). Dracula Caterpillars were found--larvae of Amblyscirtes-which have unique fangs unknown in other Lepidoptera, as well as ordinary mandibles; apparently the fangs are used in defense rather than nest-building. What was once called one species Celastrina argiolus in Colorado is now proven to be two species, with different hostplants, flight times, habitats, and pupal color; the localized species has two ecotypes, one feeding on Humulus vines, the other on Lupinus. Only one Hesperiinae (Oarisma garita) is truly polyphagous, eating grasses and sedges of many life forms; it has "satyr envy", converging to Satyrinae in its polyphagy, unique lack of larval nest, and cryptic striped larval color pattern. Hesperiinae species generally eat only a certain life form of grass, and may prefer the biochemicals of certain grass species or genera, so that Hesperiinae are comparatively host-specific. A hay-feeding guild was discovered: 6 Hesperiinae (Piruna pirus, Ancyloxypha numitor, Ochlodes sylvanoides, Poanes zabulon taxiles, Amblyscirtes vialis, probably Anatrytone logan) that each eat numerous tall wide-leaf grasses. In contrast, many Satyrinae have rather haphazard oviposition, and rather polyphagous lab feeding, so that Satyrinae species are in general rather polyphagous on various grasses or even on grasses and sedges, and their host specificity is difficult to determine. One satyr (Oeneis chryxus) oviposits on trees. In another case of convergence, Hesperia ottoe and Polites origenes are the only species in their genera to eat a broad-leaf grass and to have aerial larval nests. One skipper (Ancyloxypha) was found to have larval wax glands on four segments instead of the usual two. Bog butterflies seem to have rather polyphagous larvae. Some larvae rest underground: Parnassius (pupae), certain Satyrinae (Neominois, perhaps some Oeneis and Erebia), Hesperia relatives (Hesperia except ottoe, Polites except origenes, Yvretta, Hylephila, Atalopedes), and "Amblyscirtes" simius. Three species of Polites lay eggs without glue which drop into the litter, and Cercyonis does this about half the time. Anatrytone logan is a very distinct genus from Atrytone arogos, in contrast to Hesperia, Polites, and Atalopedes, which are basically just one genus. Cases of hostplant switching is reported in which Euphydryas chalcedona/anicia capella now feeds on introduced Linaria dalmatica, and Phyciodes picta has switched from Aster to Convolvulus. Two new subspecies are named from lowland valleys of W Colo.-E Utah: Phyciodes tharmos/morpheus riocolorado, the only valid ssp. of tharos, with paler wing color; and Hesperopsis libya confertiblanca, which has a solid white unh and a new hostplant.Item Open Access Moths of Western North America 1. Distribution of Saturniidae of Western North America(Colorado State University. Libraries, 1993) Opler, Paul A., author; Peigler, Richard S., author; C.P. Gillette Museum of Arthropod Diversity, publisherThis is the first number of a series of atlases detailing the distributional occurrence of the moths of North America. The atlas of Saturniidae by Richard Peigler and me covers the documented distribution of a well-known group.Item Open Access Atlas of western USA butterflies: including adjacent parts of Canada and Mexico(Colorado State University. Libraries, 1993-01) Stanford, Ray E., author; Opler, Paul A., authorItem Open Access Biology and systematics of Phyciodes (Phyciodes)(Colorado State University. Libraries, 1994-11-18) Scott, James A., author; James A. Scott, publisherPhyciodes (Phyciodes) is revised, using numerous new traits of larvae, pupae, hosts, larval webs, antenna color, wing pattern, male and female genitalia, etc. New hosts and life histories are presented. Twelve new names are used: five new ssp. are named (P. batesii lakota, P. b. apsaalooke, P. b. anasazi, P. pulchella shoshoni, P. p. tutchone), three names are resurrected from long disuse due to synonymy (P. pulchella = pratensis = camoestris, P. mylitta arida), homonymy (selenis vs. homonym morpheus) and synonymy (P. cocyta = selenis), two new combinations are proposed (P. pulchella montana, P. pulchella camillus), and one name is restored to species status (P. pallescens); P. vesta is removed from subgenus Phyciodes and assigned to the same subgenus (Eresia) as P. frisia. Several new western U.S. taxa proved to be ssp. of batesii based on traits of adults, larvae, pupae, diapause, hosts, and ecology. With some exceptions (antenna, some forewing traits, etc.), the tharos-group taxa form a step-cline in most traits, from P. tharos riocolorado to P. tharos to P. cocyta to B. batesii to P. pulchella; in about 10 characters, riocolorado is a "super-tharos", more extreme than tharos and thus at the end of the step-cline, while pulchella clearly forms the other end of the step-cline. Another cline appears in P. batesii. The P. mylitta-group is similar to tharos-group (mylitta/tharos share similar primitive genitalia) and contains three species that are amply distinct in larvae and male and female genitalia. Farther away, the phaon-group is newly defined by many traits of male and female genitalia and non-Aster hosts: pallescens has the wing pattern of camillus, and picta and phaon complete the group.Item Open Access Lepidoptera of North America 1. Distribution of silkmoths (Saturniidae) and hawkmoths (Sphingidae) of eastern North America(Colorado State University. Libraries, 1995) Opler, Paul A., author; Department of Entomology, Colorado State University, publisherItem Open Access Lepidoptera of North America 2. Distribution of the butterflies (Papilionoidea and Hesperioidea) of the eastern United States(Colorado State University. Libraries, 1995) Opler, Paul A., author; Department of Entomology, Colorado State University, publisherItem Open Access Moths of Western North America 2. Distribution of Sphingidae of Western North America, revised(Colorado State University. Libraries, 1995) Smith, Michael J., author; C.P. Gillette Museum of Arthropod Diversity, publisherThis is the second number of a series of atlases detailing the distributional occurrence of the moths of North America. The atlas of Sphingidae by Michael Smith covers the documented distribution of a well-known group.Item Open Access Moths of Western North America 4. Distribution of "Oecophoridae" (sense of Hodges 1983) of Western North America(Colorado State University. Libraries, 1996) Opler, Paul A., author; Powell, Jerry A., author; C.P. Gillette Museum of Arthropod Diversity, publisherThis is the fourth of a series of atlases detailing the distributional occurrence of the moths of western North America. The atlas of species traditionally assigned to "Oecophoridae" covers the documented distribution of groups of "microlepidoptera" whose taxonomic relationships are are now being reconsidered.Item Open Access Insects of Western North America 1. A survey of the Cerambycidae (Coleoptera) or longhorned beetles of Colorado(Colorado State University. Libraries, 1998) Heffern, Daniel J., author; Gillette Museum of Arthropod Diversity, Colorado State University, publisherThe purpose of this publication is to provide an account of the longhorned beetles of Colorado, to present new distributional records and to bring pertinent literature records together. One hundred ninety-three species and subspecies in 88 genera are listed, including thirty-eight new state records. The overall species distributions and host plants are included to provide an understanding of the zoogeography and possible origins of the species in the state. All available county records are included from the major institutional collections in Colorado, literature records, and numerous private collections. Previous literature citations for species not occurring or unlikely to occur in Colorado are discussed. Hyperplatys montana Casey is considered a valid species and removed from synonymy with H. aspersa (Say). Neoclytus ascendens LeConte is considered a synonym of N. leucozonus leucozonus (Castelnau & Gory).Item Open Access New western North American butterflies(Colorado State University. Libraries, 1998-02-20) Fisher, Michael S., author; Scott, James A., author; James A. Scott, publisherNew subspecies and other geographic taxa from western U.S. are described and named.Item Open Access Phyciodes (Phyciodes): new discoveries,new subspecies, and convergence(Colorado State University. Libraries, 1998-02-20) Scott, James A., author; James A. Scott, publisherThe paper presents new discoveries in Phyciodes (Phyciodes). New life history information is given for P. mylitta arizonensis, P. pallida pallida, P. pallida barnesi, P. tharos tharos, P. cocyta selenis, P. cocyta diminutor, P. batesii lakota, P. b. apsaalooke, P. b. anasazi, P. pulchella camillus, and P. picta picta. P. batesii anasazi was found to significantly converge toward P. cocyta in western Colorado in appearance of some adults larvae and pupae, and P. cocyta selenis from the same area was found to converge a little toward anasazi in adult wing pattern and larval head pattern, so some individuals of these two taxa are difficult to distinguish. The nomenclatural identity of P. tharos form marcia is clarified. Five new subspecies are named: P.tharos orantain has orange antenna nudum and is somewhat intermediate between P. tharos tharos and P. cocyta. P. cocyta diminutor is multivoltine with late-flying summer generations that are small like P. tharos, and might possibly even be a separate species from the sympatric univoltine larger P. cocyta selenis. P. pulchella deltarufa has oranger ups than ssp. pulchella. P. pulchella owimba differs from ssp. pulchella by having an orange antenna nudum and some different wing pattern details. P. phaon jalapeno has paler median ups bands.Item Open Access Speyeria hesperis and Speyeria atlantis are distinct species(Colorado State University. Libraries, 1998-02-20) Kondla, Norbert G., author; Scott, James A., author; Spomer, Stephen M., author; James A. Scott, publisherS. hesperis and S. atlantis are distinct species; past reports of "intergradation" between them actually represent polymorphism of silvering within S. hesperis. S. atlantis is always silvered, and has a darker (chocolate-brown to blackish-brown) unh disc; it occurs in eastern North America as S. a. atlantis, then ranges as S. a. hollandi over the boreal forest/aspen parklands from Man. To Alta., where it is widely sympatric with S. hesperis helena; S. a. hollandi also occurs in the Rocky Mts. in Alta.-B.C.-NE Wash.-N Ida., where it is sympatric with S. hesperis beani and S. hesperis brico (B.C., new subspecies). In the Black Hills, S. atlantis pahasapa (new subspecies) is sympatric with S. hesperis lurana. In S Wyo.-Colo.-N New Mex., S. atlantis sorocko (new subspecies) is sympatric with mostly-un silvered S. h. hesperis and mostly-silvered S. h. electa (=cornelia=nikias). S. hesperis has a redder unh disc, and ranges from Manitoba and the Black Hills westward to Alaska and the Pacific and south to New Mex.-Calif.; a majority (11 of 19) of its subspecies are usually-silvered, but only the subspecies in the extreme N and NE and S parts of its range are always silvered, and all subspecies across the middle of its range are predominantly unsilvered; silvered/unsilvered intergradation occurs within S. hesperis throughout the middle of the range, including the northern Sangre de Cristo Mts. of Colo. where unsilvered S. hesperis hesperis intergrades completely with silvered S. h. electa. Except in the extreme northern and southern ends of its range, older larvae of S. hesperis are blacker than larvae of S. atlantis.Item Open Access A new Celastrina from the eastern slope of Colorado(Colorado State University. Libraries, 1998-02-20) Wright, David M., author; Scott, James A., author; James A. Scott, publisherCelastrina humulus (new species) is named and compared to related and sympatric Celastrina. It is univoltine, and usually sympatric with the univoltine Celastrina ladon sidara and barely overlaps the end of the sidara flight, as adults emerge from post-diapause pupae later than sidara. It has a different habitat, and a different host (most populations feed on male flowers of hop Humulus lupulus, but one set of populations feeds on Lupinus argenteus flower buds). Adults are whiter than sidara. Adults are electrophoretically most similar to the eastern Prunus serotina gall feeding host race, and are somewhat similar to eastern neglecta. 1st-stage larval setae differ slightly from C. ladon sidara, mature larvae are variable but differ slightly in the frequency of color forms, and pupae differ somewhat in color, and in size of black spots.Item Open Access Supplement to Western USA/Canada/Mexico butterfly atlas: 24 February 1999 updated version(Colorado State University. Libraries, 1999) Stanford, Ray E., author; Opler, Paul A., author